Family Pycnonotidae, Bulbuls

Scaly-breasted Bulbul from Hala Bala Wildlife Sanctuary, Narathiwat, Thailand

The Bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical Asia to Indonesia, and north as far as Japan. A few insular species occur on the tropical islands of the Indian Ocean. There are 160 species in 32 genera. While different species are found in a wide range of habitats, the African species are predominantly found in rainforest, whereas Asian bulbuls are predominantly found in more open areas.

The family Pycnonotidae was introduced by the English zoologist George Robert Gray in 1840 as a subfamily Pycnonotinae of the thrush family Turdidae.

The Arabic word bulbul (بلبل) is sometimes used to refer to the "nightingale" as well as the bulbul, but the English word bulbul refers to the birds discussed in this article.

A few species that were previously considered to be members of the Pycnonotidae have been moved to other families. Several Malagasy species that were formerly placed in the genus Phyllastrephus are now placed in the family Bernieridae.[4][5] In addition, the genus Nicator containing three African species is now placed in a separate family Nicatoridae.

A study published in 2007 by Ulf Johansson and colleagues using three nuclear markers found that the genus Andropadus was non-monophyletic. In the subsequent revision, species were moved to three resurrected genera: Arizelocichla, Stelgidillas and Eurillas. Only the sombre greenbul (Andropadus importunus), was retained in Andropadus. A study by Subir Shakya and Frederick Shelden published in 2017 found that species in the large genus  Pycnonotus formed several deeply divergent clades. The genus was split and six genera were resurrected to accommodate these clades.

The family forms two main clades. One clade contains species that are only found in Africa; many of these have greenbul in the common name. The second clade contains mostly Asian species but includes a few species that are found in Africa.

The galleries below have images of the various members of Family Pycnonotidae that I have seen over the years. Simply follow the links on the images below:
 

Buff-vented Bulbul (Iole crypta) South Thailand

Baker’s Bulbul (Iole cinnamomeoventris) Kaeng Krachan

Finsch’s Bulbul (Iole finschii) Hala Bala Thailand

Grey-eyed Bulbul (Iole propinqua) Thailand

Yellow-browed Bulbul (Acritillas indica) Sri Lanka

Mountain Bulbul (Ixos mcclellandii) Thailand

Streaked Bulbul (Ixos malaccensis) Northern Thailand

Brown-eared Bulbul (Hypsipetes amaurotis) Japan

Square-tailed Black Bulbul (Hypsipetes ganeesa) Sri Lanka

Black Bulbul (Hypsipetes leucocephalus) China, Thailand

Madagascar Bulbul (Hypsipetes madagascariensis) Madagascar

Spectacled Bulbul (Pycnonotus erythropthalmos) Thailand

Scaly-breasted Bulbul (Ixodia squamata) Southern Thailand

Grey-bellied Bulbul (Ixodia cyaniventris) Thailand

Black-crested (Rubigula (Pycnonotus melanicterus) flaviventris) Thailand

Flame-throated Bulbul (Rubigula gularis) Tamil Nadu, India

Cream-vented Bulbul (Pycnonotus simplex) Southern Thailand

Red-eyed Bulbul (Pycnonotus brunneus) Southern Thailand

Streak-eared Bulbul (Pycnonotus blanfordi) Thailand

Stripe-throated (Pycnonotus finlaysoni) Thailand

Flavescent Bulbul (Pycnonotus flavescens) Thailand

Yellow-eared Bulbul (Pycnonotus penicillatus) Sri Lanka

Brown-breasted Bulbul (Pycnonotus xanthorrhous) Thailand, Yunnan

Red-whiskered Bulbul (Pycnonotus jocosus) Thailand

White-headed Bulbul (Hypsipetes thompsoni) Doi Ang Khang, Chiang Mai

Yellow-vented Bulbul (Pycnonotus analis) Thailand

Red-vented Bulbul (Pycnonotus cafer) India

Sooty-headed Bulbul (Pycnonotus aurigaster) Thailand

African Red-eyed Bulbul (Pycnonotus nigricans) Namibia

Common Bulbul (Pycnonotus barbatus) Ethiopia, East Africa

Dark-capped Bulbul (Pycnonotus tricolor) East Africa

Cape Bulbul (Pycnonotus capensis) South Africa

African Yellow Greenbul (Chlorocichla flaviventris) Namibia

Sombre Greenbul (Andropadus importunus) South Africa

Black-headed Bulbul (Pycnonotus atriceps) Thailand

Chinese Bulbul (Pycnonotus sinensis) China

Hairy-backed Bulbul (Tricholestes criniger) Southern Thailand

Ochraceus Bulbul (Alophoixus ochraceus) Thailand

Styan's Bulbul (Pycnonotus taivanus) Taipei China

Puff-throated Bulbul (Alophoixus pallidus) Thailand

Yellow-bellied Bulbul (Alophoixus phaeocephalus) Borneo

Cream-eyed Bulbul (Pycnonotus simplex) Hala Bala National Wildlife Sanctuary Thailand

Collared finchbill (Spizixos semitorques) Anhui, China

Crested Finchbill (Spizixos canifrons) Northern Thailand